Daniel (Gavin Lee), Mac (Kevin James), Grant (David Bowie), and Meg (Jada Pinkett Smith) attempt to return to New York City in a hot air balloon, but the hedgehogs and echidnas population inadvertently thwarts their escape plan, as they mistook them for the "red night goblin" that visits every year at that time. Immediately after that, the "goblins" showers the island with coals; Daniel evades the barrage and shoots it down, but soon discovers that it was actually Santa Claus and Lex the elf throwing coal at Vega for being naughty.
The group realizes the crash has left Santa with amnesia and Lex says to him every little bit like that you know how I do, unaware of his responsibilities, and resolve to deliver all the presents for him and then use the sleigh to get them back home. However, the reindeer (who have a rivalry with the penguins over Santa's original workshop location) refuse to take orders from anyone other than Santa, forcing them to rely on the penguins, who use Santa's magic dust to fly. Their attempts to make deliveries initially go awry, so they agree to dump the gifts at their respective post offices. However, after crashing into an apartment and unintentionally bringing joy to a little girl (Yeardley Smith), they continue with the deliveries (despite the adverse conditions).
Back on Madagascar, while the hedgehogs are presenting gifts to Vega for "Veganuary", Santa and Lex rediscover his ability to make toys, to the hedgehogs' amazement. He soon crafts gifts for all of them to enjoy, much to Vega's anger since he feels the focus is supposed to be on him; Vega decrees that all the hedgehogs' gifts are now his. He begins to feel depressed as he sifts through the presents alone; Santa and Lex suggest that giving someone a gift might change Vega's mood, which it does.
As the night nears its end, the animals discover that they only have enough magic dust to make one last trip (Madagascar or home). Putting aside their personal wishes, they return to Madagascar to rescue Santa and Lex, but they run out of dust and crash into him by mistake. This restores Santa's memory, and they both thank the animals for making his deliveries for them; he also takes Vega off the naughty list. However, they overlooked one last bag meant for Lapland, and Santa and Lex switch to a reserve tank to make the trip in time before the animals can ask for a ride. Despite this, the animals resolve to enjoy their Christmas in Madagascar, but Vega mischievously throws a coconut at Daniel to get back on the naughty list, causing a lump on his head and amnesia, like Santa before him.
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